Snake

Snakes are elongated, legless, carnivorous reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears. Like all squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads with their highly mobile jaws. To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca.
Living snakes are found on every continent except Antarctica, and on most smaller land masses — exceptions include some large islands, such as Ireland and New Zealand, and many small islands of the Atlantic and central Pacific. Additionally, sea snakes are widespread throughout the Indian and Pacific Oceans. More than 20 families are currently recognized, comprising about 500 genera and about 3,400 species. They range in size from the tiny, 10 cm-long thread snake to the reticulated python of up to 6.95 meters (22.8 ft) in length. The fossil species Titanoboa cerrejonensis was 13 meters (43 ft) long. Snakes are thought to have evolved from either burrowing or aquatic lizards, perhaps during the Jurassic period, with the earliest known fossils dating to between 143 and 167 Ma ago. The diversity of modern snakes appeared during the Paleocene period (c 66 to 56 Ma ago). The oldest preserved descriptions of snakes can be found in the Brooklyn Papyrus.

Most species are nonvenomous and those that have venom use it primarily to kill and subdue prey rather than for self-defense. Some possess venom potent enough to cause painful injury or death to humans. Nonvenomous snakes either swallow prey alive or kill by constriction.

>> Etymology
The English word snake comes from Old English snaca, itself from Proto-Germanic *snak-an- (cf. Germanic Schnake "ring snake", Swedish snok "grass snake"), from Proto-Indo-European root *(s)nēg-o- "to crawl", "to creep", which also gave sneak as well as Sanskrit nāgá "snake".The word ousted adder, as adder went on to narrow in meaning, though in Old English næddre was the general word for snake. The other term, serpent, is from French, ultimately from Indo-European *serp- (to creep), which also gave Ancient Greek hérpō (ἕρπω) "I crawl".

>> Evolution
The fossil record of snakes is relatively poor because snake skeletons are typically small and fragile making fossilization uncommon. Fossils readily identifiable as snakes (though often retaining hind limbs) first appear in the fossil record during the Cretaceous period. The earliest known true snake fossils (members of the crown group Serpentes) come from the marine simoliophiids, the oldest of which is the Late Cretaceous (Cenomanian age) Haasiophis terrasanctus, dated to between 112 and 94 million years old.

Based on comparative anatomy, there is consensus that snakes descended from lizards. Pythons and boas—primitive groups among modern snakes—have vestigial hind limbs: tiny, clawed digits known as anal spurs, which are used to grasp during mating. The families Leptotyphlopidae and Typhlopidae also possess remnants of the pelvic girdle, appearing as horny projections when visible.

Front limbs are nonexistent in all known snakes. This is caused by the evolution of Hox genes, controlling limb morphogenesis. The axial skeleton of the snakes’ common ancestor, like most other tetrapods, had regional specializations consisting of cervical (neck), thoracic (chest), lumbar (lower back), sacral (pelvic), and caudal (tail) vertebrae. Early in snake evolution, the Hox gene expression in the axial skeleton responsible for the development of the thorax became dominant. As a result, the vertebrae anterior to the hindlimb buds (when present) all have the same thoracic-like identity (except from the atlas, axis, and 1–3 neck vertebrae). In other words, most of a snake's skeleton is an extremely extended thorax. Ribs are found exclusively on the thoracic vertebrae. Neck, lumbar and pelvic vertebrae are very reduced in number (only 2–10 lumbar and pelvic vertebrae are present), while only a short tail remains of the caudal vertebrae. However, the tail is still long enough to be of important use in many species, and is modified in some aquatic and tree-dwelling species.

Modern snakes greatly diversified during the Paleocene. This occurred alongside the adaptive radiation of mammals, following the extinction of (non-avian) dinosaurs. The colubrids, one of the more common snake groups, became particularly diverse due to preying on rodents, an especially successful mammal group.

Origins
The origin of snakes remains an unresolved issue. There are two main hypotheses competing for acceptance.

Burrowing lizard hypothesis
There is fossil evidence to suggest that snakes may have evolved from burrowing lizards, such as the varanids (or a similar group) during the Cretaceous Period. An early fossil snake relative, Najash rionegrina, was a two-legged burrowing animal with a sacrum, and was fully terrestrial. One extant analog of these putative ancestors is the earless monitor Lanthanotus of Borneo (though it also is semiaquatic). Subterranean species evolved bodies streamlined for burrowing, and eventually lost their limbs. According to this hypothesis, features such as the transparent, fused eyelids (brille) and loss of external ears evolved to cope with fossorial difficulties, such as scratched corneas and dirt in the ears. Some primitive snakes are known to have possessed hindlimbs, but their pelvic bones lacked a direct connection to the vertebrae. These include fossil species like Haasiophis, Pachyrhachis and Eupodophis, which are slightly older than Najash.
This hypothesis was strengthened in 2015 by the discovery of a 113m year-old fossil of a four-legged snake in Brazil that has been named Tetrapodophis amplectus. It has many snake-like features, is adapted for burrowing and its stomach indicates that it was preying on other animals.It is currently uncertain if Tetrapodophis is a snake or another species, in the squamate order, as a snake-like body has independently evolved at least 26 times. Tetrapodophis does not have distinctive snake features in its spine and skull.

Aquatic mosasaur hypothesis
An alternative hypothesis, based on morphology, suggests the ancestors of snakes were related to mosasaurs—extinct aquatic reptiles from the Cretaceous—which in turn are thought to have derived from varanid lizards.According to this hypothesis, the fused, transparent eyelids of snakes are thought to have evolved to combat marine conditions (corneal water loss through osmosis), and the external ears were lost through disuse in an aquatic environment. This ultimately lead to an animal similar to today's sea snakes. In the Late Cretaceous, snakes recolonized land, and continued to diversify into today's snakes. Fossilized snake remains are known from early Late Cretaceous marine sediments, which is consistent with this hypothesis; particularly so, as they are older than the terrestrial Najash rionegrina. Similar skull structure, reduced or absent limbs, and other anatomical features found in both mosasaurs and snakes lead to a positive cladistical correlation, although some of these features are shared with varanids.

Genetic studies in recent years have indicated snakes are not as closely related to monitor lizards as was once believed—and therefore not to mosasaurs, the proposed ancestor in the aquatic scenario of their evolution. However, more evidence links mosasaurs to snakes than to varanids. Fragmented remains found from the Jurassic and Early Cretaceous indicate deeper fossil records for these groups, which may potentially refute either hypothesis.

>> Distribution
There are over 2,900 species of snakes ranging as far northward as the Arctic Circle in Scandinavia and southward through Australia. Snakes can be found on every continent except Antarctica, in the sea, and as high as 16,000 feet (4,900 m) in the Himalayan Mountains of Asia. There are numerous islands from which snakes are absent, such as Ireland, Iceland, and New Zealand (although New Zealand's waters are infrequently visited by the yellow-bellied sea snake and the banded sea krait).

>> Taxonomy
See also: List of snake genera
All modern snakes are grouped within the suborder Serpentes in Linnean taxonomy, part of the order Squamata, though their precise placement within squamates remains controversial.

The two infraorders of Serpentes are: Alethinophidia and Scolecophidia. This separation is based on morphological characteristics and mitochondrial DNA sequence similarity. Alethinophidia is sometimes split into Henophidia and Caenophidia, with the latter consisting of "colubroid" snakes (colubrids, vipers, elapids, hydrophiids, and atractaspids) and acrochordids, while the other alethinophidian families comprise Henophidia.While not extant today, the Madtsoiidae, a family of giant, primitive, python-like snakes, was around until 50,000 years ago in Australia, represented by genera such as Wonambi.

There are numerous debates in the systematics within the group. For instance, many sources classify Boidae and Pythonidae as one family, while some keep the Elapidae and Hydrophiidae (sea snakes) separate for practical reasons despite their extremely close relation.

Recent molecular studies support the monophyly of the clades of modern snakes, scolecophidians, typhlopids + anomalepidids, alethinophidians, core alethinophidians, uropeltids (Cylindrophis, Anomochilus, uropeltines), macrostomatans, booids, boids, pythonids and caenophidians

>> Biology
> Size
The now extinct Titanoboa cerrejonensis snakes found were 12.8–13 m (42–43 ft) in length. By comparison, the largest extant snakes are the reticulated python, the longest recorded specimen measured about 6.95 m (22.8 ft) long, and the anaconda, which measures about 5.21 m (17.1 ft) long and is considered the heaviest snake on Earth.

At the other end of the scale, the smallest extant snake is Leptotyphlops carlae, with a length of about 10 cm (3.9 in).Most snakes are fairly small animals, approximately 1 m (3.3 ft) in length.

> Perception
Smell
Snakes use smell to track their prey. They smell by using their forked tongues to collect airborne particles, then passing them to the vomeronasal organ or Jacobson's organ in the mouth for examination.The fork in the tongue gives snakes a sort of directional sense of smell and taste simultaneously. They keep their tongues constantly in motion, sampling particles from the air, ground, and water, analyzing the chemicals found, and determining the presence of prey or predators in the local environment. In water-dwelling snakes, such as the anaconda, the tongue functions efficiently under water.
Eyesight
Snake vision varies greatly, from only being able to distinguish light from dark to keen eyesight, but the main trend is that their vision is adequate although not sharp, and allows them to track movements. Generally, vision is best in arboreal snakes and weakest in burrowing snakes. Some snakes, such as the Asian vine snake (genus Ahaetulla), have binocular vision, with both eyes capable of focusing on the same point. Most snakes focus by moving the lens back and forth in relation to the retina, while in the other amniote groups, the lens is stretched. Many nocturnal snakes have slit pupils while diurnal snakes have round pupils.
Infrared sensitivity
Pit vipers, pythons, and some boas have infrared-sensitive receptors in deep grooves on the snout, which allow them to "see" the radiated heat of warm-blooded prey mammals. In pit vipers the grooves are located between the nostril and the eye, in a large "pit" on each side of the head. Other infrared-sensitive snakes have multiple, smaller labial pits lining the upper lip, just below the nostrils.
Vibration sensitivity
The part of the body in direct contact with the ground is very sensitive to vibration; thus, a snake can sense other animals approaching by detecting faint vibrations in the air and on the ground
Skin
Main article: Snake scales
The skin of a snake is covered in scales. Contrary to the popular notion of snakes being slimy because of possible confusion of snakes with worms, snakeskin has a smooth, dry texture. Most snakes use specialized belly scales to travel, gripping surfaces. The body scales may be smooth, keeled, or granular. The eyelids of a snake are transparent "spectacle" scales, which remain permanently closed, also known as brille.

The shedding of scales is called ecdysis (or in normal usage, molting or sloughing). In the case of snakes, the complete outer layer of skin is shed in one layer. Snake scales are not discrete, but extensions of the epidermis—hence they are not shed separately but as a complete outer layer during each molt, akin to a sock being turned inside out.

The shape and number of scales on the head, back, and belly are often characteristic and used for taxonomic purposes. Scales are named mainly according to their positions on the body. In "advanced" (Caenophidian) snakes, the broad belly scales and rows of dorsal scales correspond to the vertebrae, allowing scientists to count the vertebrae without dissection.


Eye scales visible during the molt of a diamond python.
Snakes' eyes are covered by their clear scales (the brille) rather than movable eyelids. Their eyes are always open, and for sleeping, the retina can be closed or the face buried among the folds of the body.

Moulting
Moulting serves a number of functions. Firstly, the old and worn skin is replaced; secondly, it helps get rid of parasites such as mites and ticks. Renewal of the skin by moulting is supposed to allow growth in some animals such as insects; however, this has been disputed in the case of snakes.
Molting occurs periodically throughout the snake's life. Before a molt, the snake stops eating and often hides or moves to a safe place. Just before shedding, the skin becomes dull and dry looking and the eyes become cloudy or blue-colored. The inner surface of the old skin liquefies. This causes the old skin to separate from the new skin beneath it. After a few days, the eyes clear and the snake "crawls" out of its old skin. The old skin breaks near the mouth and the snake wriggles out, aided by rubbing against rough surfaces. In many cases, the cast skin peels backward over the body from head to tail in one piece, like pulling a sock off inside-out. A new, larger, brighter layer of skin has formed underneath.
The discarded skin gives a perfect imprint of the scale pattern, and it is usually possible to identify the snake if the discarded skin is reasonably intact. This periodic renewal has led to the snake being a symbol of healing and medicine, as pictured in the Rod of Asclepius.

Skeleton
The skeleton of most snakes consists solely of the skull, hyoid, vertebral column, and ribs, though henophidian snakes retain vestiges of the pelvis and rear limbs.

The skull of the snake consists of a solid and complete neurocranium, to which many of the other bones are only loosely attached, particularly the highly mobile jaw bones, which facilitate manipulation and ingestion of large prey items. The left and right sides of the lower jaw are joined only by a flexible ligament at the anterior tips, allowing them to separate widely, while the posterior end of the lower jaw bones articulate with a quadrate bone, allowing further mobility. The bones of the mandible and quadrate bones can also pick up ground borne vibrations. Because the sides of the jaw can move independently of one another, snakes resting their jaws on a surface have sensitive stereo hearing which can detect the position of prey. The jaw-quadrate-stapes pathway is capable of detecting vibrations on the angstrom scale, despite the absence of an outer ear and the ossicle mechanism of impedance matching used in other vertebrates to receive vibrations from the air.

The hyoid is a small bone located posterior and ventral to the skull, in the 'neck' region, which serves as an attachment for muscles of the snake's tongue, as it does in all other tetrapods.

The vertebral column consists of anywhere between 200 to 400 (or more) vertebrae. Tail vertebrae are comparatively few in number (often less than 20% of the total) and lack ribs, while body vertebrae each have two ribs articulating with them. The vertebrae have projections that allow for strong muscle attachment enabling locomotion without limbs.

Autotomy of the tail, a feature found in some lizards is absent in most snakes.Caudal autotomy in snakes is rare and is intervertebral, unlike that in lizards, which is intravertebral—that is, the break happens along a predefined fracture plane present on a vertebra.

In some snakes, most notably boas and pythons, there are vestiges of the hindlimbs in the form of a pair of pelvic spurs. These small, claw-like protrusions on each side of the cloaca are the external portion of the vestigial hindlimb skeleton, which includes the remains of an ilium and femur.

Internal organs
The snake's heart is encased in a sac, called the pericardium, located at the bifurcation of the bronchi. The heart is able to move around, however, owing to the lack of a diaphragm. This adjustment protects the heart from potential damage when large ingested prey is passed through the esophagus. The spleen is attached to the gall bladder and pancreas and filters the blood. The thymus gland is located in fatty tissue above the heart and is responsible for the generation of immune cells in the blood. The cardiovascular system of snakes is also unique for the presence of a renal portal system in which the blood from the snake's tail passes through the kidneys before returning to the heart.

The vestigial left lung is often small or sometimes even absent, as snakes' tubular bodies require all of their organs to be long and thin. In the majority of species, only one lung is functional. This lung contains a vascularized anterior portion and a posterior portion that does not function in gas exchange. This 'saccular lung' is used for hydrostatic purposes to adjust buoyancy in some aquatic snakes and its function remains unknown in terrestrial species.Many organs that are paired, such as kidneys or reproductive organs, are staggered within the body, with one located ahead of the other.

Snakes have no lymph nodes

>> Venom
Cobras, vipers, and closely related species use venom to immobilize or kill their prey. The venom is modified saliva, delivered through fangs. The fangs of 'advanced' venomous snakes like viperids and elapids are hollow to inject venom more effectively, while the fangs of rear-fanged snakes such as the boomslang merely have a groove on the posterior edge to channel venom into the wound. Snake venoms are often prey specific—their role in self-defense is secondary.

Venom, like all salivary secretions, is a predigestant that initiates the breakdown of food into soluble compounds, facilitating proper digestion. Even nonvenomous snake bites (like any animal bite) will cause tissue damage.

Certain birds, mammals, and other snakes (such as kingsnakes) that prey on venomous snakes have developed resistance and even immunity to certain venoms. Venomous snakes include three families of snakes, and do not constitute a formal classification group used in taxonomy.

The term "poisonous snake" is mostly incorrect. Poison is inhaled or ingested, whereas venom is injected.There are, however, two exceptions: Rhabdophis sequesters toxins from the toads it eats, then secretes them from nuchal glands to ward off predators, and a small population of garter snakes in Oregon retains enough toxin in their liver from the newts they eat to be effectively poisonous to small local predators (such as crows and foxes).

Snake venoms are complex mixtures of proteins, and are stored in venom glands at the back of the head.In all venomous snakes, these glands open through ducts into grooved or hollow teeth in the upper jaw.These proteins can potentially be a mix of neurotoxins (which attack the nervous system), hemotoxins (which attack the circulatory system), cytotoxins, bungarotoxins and many other toxins that affect the body in different ways.Almost all snake venom contains hyaluronidase, an enzyme that ensures rapid diffusion of the venom

Venomous snakes that use hemotoxins usually have fangs in the front of their mouths, making it easier for them to inject the venom into their victims.Some snakes that use neurotoxins (such as the mangrove snake) have fangs in the back of their mouths, with the fangs curled backwards.This makes it difficult both for the snake to use its venom and for scientists to milk them.Elapids, however, such as cobras and kraits are proteroglyphous—they possess hollow fangs that cannot be erected toward the front of their mouths, and cannot "stab" like a viper. They must actually bite the victim.

It has recently been suggested that all snakes may be venomous to a certain degree, with harmless snakes having weak venom and no fangs. Most snakes currently labelled "nonvenomous" would still be considered harmless according to this theory, as they either lack a venom delivery method or are incapable of delivering enough to endanger a human. This theory postulates that snakes may have evolved from a common lizard ancestor that was venomous—and that venomous lizards like the gila monster, beaded lizard, monitor lizards, and the now-extinct mosasaurs may also have derived from it. They share this venom clade with various other saurian species.

Venomous snakes are classified in two taxonomic families:

Elapids – cobras including king cobras, kraits, mambas, Australian copperheads, sea snakes, and coral snakes.
Viperids – vipers, rattlesnakes, copperheads/cottonmouths, and bushmasters.
There is a third family containing the opistoglyphous (rear-fanged) snakes (as well as the majority of other snake species):

Colubrids – boomslangs, tree snakes, vine snakes, mangrove snakes, although not all colubrids are venomous.
Reproduction
See also: Sexual selection in scaled reptiles
Although a wide range of reproductive modes are used by snakes, all snakes employ internal fertilization. This is accomplished by means of paired, forked hemipenes, which are stored, inverted, in the male's tail. The hemipenes are often grooved, hooked, or spined in order to grip the walls of the female's cloaca.

Most species of snakes lay eggs, but most snakes abandon the eggs shortly after laying. However, a few species (such as the king cobra) actually construct nests and stay in the vicinity of the hatchlings after incubation. Most pythons coil around their egg-clutches and remain with them until they hatch. A female python will not leave the eggs, except to occasionally bask in the sun or drink water. She will even "shiver" to generate heat to incubate the eggs.

Some species of snake are ovoviviparous and retain the eggs within their bodies until they are almost ready to hatch. Recently, it has been confirmed that several species of snake are fully viviparous, such as the boa constrictor and green anaconda, nourishing their young through a placenta as well as a yolk sac, which is highly unusual among reptiles, or anything else outside of requiem sharks or placental mammals. Retention of eggs and live birth are most often associated with colder environments.
Sexual selection in snakes is demonstrated by the three thousand species each use different tactics in acquiring mates.Ritual combat between males for the females they want to mate with includes topping, a behavior exhibited by most viperids in which one male will twist around the vertically elevated fore body of its opponent and forcing it downward. It is common for neck biting to occur while the snakes are entwined.

Behavior
Winter dormancy
In regions where winters are colder than snakes can tolerate while remaining active, local species will brumate. Unlike hibernation, in which mammals are actually asleep, brumating reptiles are awake but inactive. Individual snakes may brumate in burrows, under rock piles, or inside fallen trees, or snakes may aggregate in large numbers at hibernacula.

Feeding and diet
All snakes are strictly carnivorous, eating small animals including lizards, frogs, other snakes, small mammals, birds, eggs, fish, snails or insects. Because snakes cannot bite or tear their food to pieces, they must swallow prey whole. The body size of a snake has a major influence on its eating habits. Smaller snakes eat smaller prey. Juvenile pythons might start out feeding on lizards or mice and graduate to small deer or antelope as an adult, for example.

The snake's jaw is a complex structure. Contrary to the popular belief that snakes can dislocate their jaws, snakes have a very flexible lower jaw, the two halves of which are not rigidly attached, and numerous other joints in their skull (see snake skull), allowing them to open their mouths wide enough to swallow their prey whole, even if it is larger in diameter than the snake itself. For example, the African egg-eating snake has flexible jaws adapted for eating eggs much larger than the diameter of its head.This snake has no teeth, but does have bony protrusions on the inside edge of its spine, which it uses to break shells when it eats eggs.

While the majority of snakes eat a variety of prey animals, there is some specialization by some species. King cobras and the Australian bandy-bandy consume other snakes. Pareas iwesakii and other snail-eating colubrids of subfamily Pareatinae have more teeth on the right side of their mouths than on the left, as the shells of their prey usually spiral clockwise

Some snakes have a venomous bite, which they use to kill their prey before eating it. Other snakes kill their prey by constriction. Still others swallow their prey whole and alive.

After eating, snakes become dormant while the process of digestion takes place. Digestion is an intense activity, especially after consumption of large prey. In species that feed only sporadically, the entire intestine enters a reduced state between meals to conserve energy. The digestive system is then 'up-regulated' to full capacity within 48 hours of prey consumption. Being ectothermic ("cold-blooded"), the surrounding temperature plays a large role in snake digestion. The ideal temperature for snakes to digest is 30 °C (86 °F). So much metabolic energy is involved in a snake's digestion that in the Mexican rattlesnake (Crotalus durissus), surface body temperature increases by as much as 1.2 °C (2.2 °F) during the digestive process.Because of this, a snake disturbed after having eaten recently will often regurgitate its prey to be able to escape the perceived threat. When undisturbed, the digestive process is highly efficient, with the snake's digestive enzymes dissolving and absorbing everything but the prey's hair (or feathers) and claws, which are excreted along with waste.

Locomotion
The lack of limbs does not impede the movement of snakes. They have developed several different modes of locomotion to deal with particular environments. Unlike the gaits of limbed animals, which form a continuum, each mode of snake locomotion is discrete and distinct from the others; transitions between modes are abrupt.

Lateral undulation
Main article: Undulatory locomotion
Lateral undulation is the sole mode of aquatic locomotion, and the most common mode of terrestrial locomotion.In this mode, the body of the snake alternately flexes to the left and right, resulting in a series of rearward-moving "waves". While this movement appears rapid, snakes have rarely been documented moving faster than two body-lengths per second, often much less. This mode of movement has the same net cost of transport (calories burned per meter moved) as running in lizards of the same mass.

Terrestrial
Terrestrial lateral undulation is the most common mode of terrestrial locomotion for most snake species. In this mode, the posteriorly moving waves push against contact points in the environment, such as rocks, twigs, irregularities in the soil, etc. Each of these environmental objects, in turn, generates a reaction force directed forward and towards the midline of the snake, resulting in forward thrust while the lateral components cancel out. The speed of this movement depends upon the density of push-points in the environment, with a medium density of about 8 along the snake's length being ideal.The wave speed is precisely the same as the snake speed, and as a result, every point on the snake's body follows the path of the point ahead of it, allowing snakes to move through very dense vegetation and small openings.

Monkey

Monkeys are haplorhine ("dry-nosed") primates, a paraphyletic group generally possessing tails and consisting of approximately 260 known living species. Many monkey species are tree-dwelling (arboreal), although there are species that live primarily on the ground, such as baboons. Most species are also active during the day (diurnal). Monkeys are generally considered to be intelligent, particularly Old World monkeys.

Lemurs, lorises, and galagos are not monkeys; instead they are strepsirrhine ("wet-nosed") primates. Like monkeys, tarsiers are haplorhine primates; however, they are also not monkeys. There are two major types of monkey: New World monkeys (platyrrhines) from South and Central America and Old World monkeys (catarrhines of the superfamily Cercopithecoidea) from Africa and Asia. Hominoid apes (gibbons, orangutans, gorillas, chimpanzees, and humans), which all lack tails, are also catarrhines but are not considered monkeys.(Tailless monkeys may be called "apes", incorrectly according to modern usage; thus the tailless Barbary macaque is sometimes called the "Barbary ape".) Because old world monkeys are more closely related to hominoid apes than to new world monkeys, yet the term "monkey" excludes these closer relatives, monkeys are referred to as a paraphyletic group. Simians ("monkeys") and tarsiers emerged within haplorrhines some 60 million years ago. New world monkeys and catarrhine monkeys emerged within the simians some 35 millions years ago. Old world monkeys and Hominoidea emerged within the catarrhine monkeys some 25 millions years ago. Extinct basal simians such as Aegyptopithecus or Parapithecus are also considered monkeys by primatologists.

>> Historical and modern terminology
The origin of the word is unclear, however, it is likely derived from Sanskrit word "markata", which denominates non-primate apes. In most Germanic languages monkeys are called "meercat" in accordance to that.(In English the meaning of meercat has changed to denominate a different animal.) According to the Online Etymology Dictionary, the word "monkey" may originate in a German version of the Reynard the Fox fable, published circa 1580. In this version of the fable, a character named Moneke is the son of Martin the Ape. In English, no very clear distinction was originally made between "ape" and "monkey"; thus the 1910 Encyclopædia Britannica entry for "ape" notes that it is either a synonym for "monkey" or is used to mean a tailless humanlike primate. Colloquially, the terms "monkey" and "ape" are widely used interchangeably. Also, a few monkey species have the word "ape" in their common name, such as the Barbary ape.

Later in the first half of the 20th century, the idea developed that there were trends in primate evolution and that the living members of the order could be arranged in a series, leading through "monkeys" and "apes" to humans. Monkeys thus constituted a "grade" on the path to humans and were distinguished from "apes".

Scientific classifications are now more often based on monophyletic groups, that is groups consisting of all the descendants of a common ancestor. The New World monkeys and the Old World monkeys are each monophyletic groups, but their combination is not, since it excludes hominoids (apes and humans). Thus the term "monkey" no longer refers to a recognized scientific taxon. The smallest accepted taxon which contains all the monkeys is the infraorder Simiiformes, or simians. However this also contains the hominoids (apes and humans), so that monkeys are, in terms of currently recognized taxa, non-hominoid simians. Colloquially and pop-culturally, the term is ambiguous and sometimes monkey includes non-human hominoids.

A group of monkeys may be commonly referred to as a tribe or a troop

>> Description
Monkeys range in size from the pygmy marmoset, which can be as small as 117 millimetres (4.6 in) with a 172-millimetre (6.8 in) tail and just over 100 grams (3.5 oz) in weight,to the male mandrill, almost 1 metre (3.3 ft) long and weighing up to 36 kilograms (79 lb).Some are arboreal (living in trees) while others live on the savanna; diets differ among the various species but may contain any of the following: fruit, leaves, seeds, nuts, flowers, eggs and small animals (including insects and spiders).

Some characteristics are shared among the groups; most New World monkeys have prehensile tails while Old World monkeys have non-prehensile tails or no visible tail at all. Old World monkeys have trichromatic color vision like that of humans, while New World monkeys may be trichromatic, dichromatic, or—as in the owl monkeys and greater galagos—monochromatic. Although both the New and Old World monkeys, like the apes, have forward-facing eyes, the faces of Old World and New World monkeys look very different, though again, each group shares some features such as the types of noses, cheeks and rumps.

>> Classification
The following list shows where the various monkey families (bolded) are placed in the classification of living (extant) primates.

ORDER PRIMATES
Suborder Strepsirrhini: lemurs, lorises, and galagos
Suborder Haplorhini: tarsiers, monkeys, and apes
Infraorder Tarsiiformes
Family Tarsiidae: tarsiers
Infraorder Simiiformes: simians
Parvorder Platyrrhini: New World monkeys
Family Callitrichidae: marmosets and tamarins (42 species)
Family Cebidae: capuchins and squirrel monkeys (14 species)
Family Aotidae: night monkeys (11 species)
Family Pitheciidae: titis, sakis, and uakaris (41 species)
Family Atelidae: howler, spider, and woolly monkeys (24 species)
Parvorder Catarrhini
Superfamily Cercopithecoidea
Family Cercopithecidae: Old World monkeys (135 species)
Superfamily Hominoidea: apes
Family Hylobatidae: gibbons ("lesser apes") (17 species)
Family Hominidae: great apes including humans (7 species)

>> Relationship with humans
The many species of monkey have varied relationships with humans. Some are kept as pets, others used as model organisms in laboratories or in space missions. They may be killed in monkey drives (when they threaten agriculture) or used as service animals for the disabled.

In some areas, some species of monkey are considered agricultural pests, and can cause extensive damage to commercial and subsistence crops. This can have important implications for the conservation of endangered species, which may be subject to persecution. In some instances farmers' perceptions of the damage may exceed the actual damage.Monkeys that have become habituated to human presence in tourist locations may also be considered pests, attacking tourists.

In religion and culture, the monkey often represents quick-wittedness and mischief.

As service animals for the disabled
Some organizations train capuchin monkeys as service animals to assist quadriplegics and other people with severe spinal cord injuries or mobility impairments. After being socialized in a human home as infants, the monkeys undergo extensive training before being placed with a disabled person. Around the house, the monkeys assist with feeding, fetching, manipulating objects, and personal care.

In experiments
The most common monkey species found in animal research are the grivet, the rhesus macaque, and the crab-eating macaque, which are either wild-caught or purpose-bred.They are used primarily because of their relative ease of handling, their fast reproductive cycle (compared to apes) and their psychological and physical similarity to humans. Worldwide, it is thought that between 100,000 and 200,000 non-human primates are used in research each year,64.7% of which are Old World monkeys, and 5.5% New World monkeys. This number makes a very small fraction of all animals used in research.Between 1994 and 2004 the United States has used an average of 54,000 non-human primates, while around 10,000 non-human primates were used in the European Union in 2002.
The use of monkeys in laboratories is controversial. Animal rights activists claim that their use is cruel and produces little information of value, and there have been many protests, vandalism to testing facilities and threats to workers. Others claim that it has led to many important medical breakthroughs such as the rabies vaccine, understanding of human reproduction and basic knowledge about brain function and that the prevention of harm to humans should be a higher priority than the possible harm done to monkeys. The topic has become a popular cause for animal rights and animal welfare groups.

In space
Main article: Monkeys in space
A number of countries have used monkeys as part of their space exploration programmes, including the United States and France. The first monkey in space was Albert II, who flew in the US-launched V-2 rocket on June 14, 1949.

As food
Monkey brains are eaten as a delicacy in parts of South Asia, Africa and China.In traditional Islamic dietary laws, the eating of monkeys is forbidden. However, monkeys are sometimes eaten in parts of Africa, where they can be sold as "bushmeat".

Literature
Sun Wukong (the "Monkey King"), a character who figures prominently in Chinese mythology, is the protagonist in the classic comic Chinese novel Journey to the West.

Monkeys are prevalent in numerous books, television programs, and movies. The television series Monkey and the literary characters Monsieur Eek and Curious George are all examples.

Informally, the term "monkey" is often used more broadly than in scientific use, and may be used to refer to apes, particularly chimpanzees, gibbons, and gorillas. Author Terry Pratchett alludes to this difference in usage in his Discworld novels, in which the Librarian of the Unseen University is an orangutan who gets very violent if referred to as a monkey. Another example is the use of Simians in Chinese poetry.

The Winged monkeys are prominent characters in The Wizard of Oz.

Religion and worship
Hanuman, a prominent divine entity in Hinduism, is a Human-like monkey god. He bestows courage, strength and longevity to the person who thinks about him or the god Rama.

In Buddhism, the monkey is an early incarnation of Buddha but may also represent trickery and ugliness. The Chinese Buddhist "mind monkey" metaphor refers to the unsettled, restless state of human mind. Monkey is also one of the Three Senseless Creatures, symbolizing greed, with the tiger representing anger and the deer lovesickness.

The Mizaru, or three wise monkeys, are revered in Japanese folklore, together they embody the proverbial principle to "see no evil, hear no evil, speak no evil".

The Moche people of ancient Peru worshipped nature.They placed emphasis on animals and often depicted monkeys in their art.

The Tzeltal people of Mexico worshipped monkeys as incarnations of their dead ancestors.

Zodiac
The Monkey (猴) is the ninth in the twelve-year cycle of animals which appear in the Chinese zodiac related to the Chinese calendar. Next year will be the year of the monkey

Chicken

The chicken (Gallus gallus domesticus) is a domesticated fowl, a subspecies of the red junglefowl. As one of the most common and widespread domestic animals, with a population of more than 24 billion in 2003, there are more chickens in the world than any other species of bird. Humans keep chickens primarily as a source of food, consuming both their meat and their eggs.

The earliest known domestication occurred in Northern China 8,000 BCE determined from archaeological chicken bones. Previously genetic studies have pointed to multiple maternal origins in Southeast, East, and South Asia, but with the clade found in the Americas, Europe, the Middle East and Africa originating in the Indian subcontinent. From India, the domesticated chicken was imported to Lydia in western Asia Minor, and to Greece by the fifth century BC.Fowl had been known in Egypt since the mid-15th century BC, with the "bird that gives birth every day" having come to Egypt from the land between Syria and Shinar, Babylonia, according to the annals of Thutmose III.

>> Terminology
In the UK and Ireland adult male chickens over the age of one year are primarily known as cocks, whereas in America, Australia and Canada they are more commonly called roosters. Males less than a year old are cockerels. Castrated roosters are called capons (surgical and chemical castration are now illegal in some parts of the world). Females over a year old are known as hens and younger females as pullets although in the egg-laying industry, a pullet becomes a hen when she begins to lay eggs at 16 to 20 weeks of age. In Australia and New Zealand (also sometimes in Britain), there is a generic term chook /ˈtʃʊk/ to describe all ages and both sexes. The young are called chicks and the meat is called chicken.

"Chicken" originally referred to chicks, not the species itself. The species as a whole was then called domestic fowl, or just fowl. This use of "chicken" survives in the phrase "Hen and Chickens", sometimes used as a British public house or theatre name, and to name groups of one large and many small rocks or islands in the sea (see for example Hen and Chicken Islands). The word "chicken" is sometimes erroneously construed to mean females exclusively, despite the term "hen" for females being in wide circulation.

In the Deep South of the United States chickens are also referred to by the slang term yardbird.

>> General biology and habitat
Chickens are omnivores. In the wild, they often scratch at the soil to search for seeds, insects and even larger animals such as lizards, small snakes or young mice.

Chickens may live for five to ten years, depending on the breed. The world's oldest chicken, a hen, died of heart failure at the age of 16 according to Guinness World Records.

Roosters can usually be differentiated from hens by their striking plumage of long flowing tails and shiny, pointed feathers on their necks (hackles) and backs (saddle), which are typically of brighter, bolder colours than those of females of the same breed. However, in some breeds, such as the Sebright chicken, the rooster has only slightly pointed neck feathers, the same colour as the hen's. The identification can be made by looking at the comb, or eventually from the development of spurs on the male's legs (in a few breeds and in certain hybrids, the male and female chicks may be differentiated by colour). Adult chickens have a fleshy crest on their heads called a comb, or cockscomb, and hanging flaps of skin either side under their beaks called wattles. Collectively, these and other fleshy protuberances on the head and throat are called caruncles. Both the adult male and female have wattles and combs, but in most breeds these are more prominent in males. A muff or beard is a mutation found in several chicken breeds which causes extra feathering under the chicken's face, giving the appearance of a beard. Domestic chickens are not capable of long distance flight, although lighter birds are generally capable of flying for short distances, such as over fences or into trees (where they would naturally roost). Chickens may occasionally fly briefly to explore their surroundings, but generally do so only to flee perceived danger.

>> Behaviour
Social behaviour
Chickens are gregarious birds and live together in flocks. They have a communal approach to the incubation of eggs and raising of young. Individual chickens in a flock will dominate others, establishing a "pecking order", with dominant individuals having priority for food access and nesting locations. Removing hens or roosters from a flock causes a temporary disruption to this social order until a new pecking order is established. Adding hens, especially younger birds, to an existing flock can lead to fighting and injury.When a rooster finds food, he may call other chickens to eat first. He does this by clucking in a high pitch as well as picking up and dropping the food. This behaviour may also be observed in mother hens to call their chicks and encourage them to eat.

A rooster's crowing (a loud and sometimes shrill call) is a territorial signal to other roosters. However, crowing may also result from sudden disturbances within their surroundings. Hens cluck loudly after laying an egg, and also to call their chicks. Chickens also give a low "warning call" when they think they see a predator approaching.

Courtship
To initiate courting, some roosters may dance in a circle around or near a hen ("a circle dance"), often lowering his wing which is closest to the hen.The dance triggers a response in the hen and when she responds to his "call", the rooster may mount the hen and proceed with the mating.

>> Nesting and laying behaviour
Hens will often try to lay in nests that already contain eggs and have been known to move eggs from neighbouring nests into their own. The result of this behaviour is that a flock will use only a few preferred locations, rather than having a different nest for every bird. Hens will often express a preference to lay in the same location. It is not unknown for two (or more) hens to try to share the same nest at the same time. If the nest is small, or one of the hens is particularly determined, this may result in chickens trying to lay on top of each other. There is evidence that individual hens prefer to be either solitary or gregarious nesters.Some farmers use fake eggs made from plastic or stone (or golf balls) to encourage hens to lay in a particular location.

Broodiness
Under natural conditions, most birds lay only until a clutch is complete, and they will then incubate all the eggs. Many domestic hens will also do this–and are then said to "go broody". The broody hen will stop laying and instead will focus on the incubation of the eggs (a full clutch is usually about 12 eggs). She will "sit" or "set" on the nest, protesting or pecking in defense if disturbed or removed, and she will rarely leave the nest to eat, drink, or dust-bathe. While brooding, the hen maintains the nest at a constant temperature and humidity, as well as turning the eggs regularly during the first part of the incubation. To stimulate broodiness, an owner may place many artificial eggs in the nest, or to stop it they may place the hen in an elevated cage with an open wire floor.
Modern egg-laying breeds rarely go broody, and those that do often stop part-way through the incubation. However, some "utility" (general purpose) breeds, such as the Cochin, Cornish and Silkie, do regularly go broody, and they make excellent mothers, not only for chicken eggs but also for those of other species—even those with much smaller or larger eggs and different incubation periods, such as quail, pheasants, turkeys or geese. Chicken eggs can also be hatched under a broody duck, with varied success.

Hatching and early life
At the end of the incubation period (about 21 days),the eggs, if fertile, will hatch. Development of the egg starts only when incubation begins, so they all hatch within a day or two of each other, despite perhaps being laid over a period of two weeks or so. Before hatching, the hen can hear the chicks peeping inside the eggs, and will gently cluck to stimulate them to break out of their shells. The chick begins by "pipping"; pecking a breathing hole with its egg tooth towards the blunt end of the egg, usually on the upper side. The chick will then rest for some hours, absorbing the remaining egg yolk and withdrawing the blood supply from the membrane beneath the shell (used earlier for breathing through the shell). It then enlarges the hole, gradually turning round as it goes, and eventually severing the blunt end of the shell completely to make a lid. It crawls out of the remaining shell, and its wet down dries out in the warmth of the nest.

The hen will usually stay on the nest for about two days after the first egg hatches, and during this time the newly hatched chicks live off the egg yolk they absorb just before hatching. Any eggs not fertilized by a rooster will not hatch, and the hen eventually loses interest in these and leaves the nest. After hatching, the hen fiercely guards the chicks, and will brood them when necessary to keep them warm, at first often returning to the nest at night. She leads them to food and water; she will call them to edible items, but seldom feeds them directly. She continues to care for them until they are several weeks old, when she will gradually lose interest and eventually start to lay again.

>> Embryology
In 2006, scientists researching the ancestry of birds "turned on" a chicken recessive gene, talpid2, and found that the embryo jaws initiated formation of teeth, like those found in ancient bird fossils. John Fallon, the overseer of the project, stated that chickens have "...retained the ability to make teeth, under certain conditions... ."

>> Breeding
Origins

The domestic chicken is descended primarily from the red junglefowl (Gallus gallus) and is scientifically classified as the same species. As such it can and does freely interbreed with populations of red jungle fowl. Recent genetic analysis has revealed that at least the gene for yellow skin was incorporated into domestic birds through hybridization with the grey junglefowl (G. sonneratii). The traditional poultry farming view is stated in Encyclopædia Britannica (2007): "Humans first domesticated chickens of Indian origin for the purpose of cockfighting in Asia, Africa, and Europe. Very little formal attention was given to egg or meat production... ", a view supported by many archeologists. In the last decade there have been a number of genetic studies. According to one study, a single domestication event occurring in the region of modern Thailand created the modern chicken with minor transitions separating the modern breeds.However, that study was later found to be based on incomplete data, and recent studies point to multiple maternal origins, with the clade found in the Americas, Europe, Middle East, and Africa, originating from the Indian subcontinent, where a large number of unique haplotypes occur. It is postulated that the jungle fowl, known as the "bamboo fowl" in many Southeast Asian languages, is a special pheasant well adapted to take advantage of the large amounts of fruits that are produced during the end of the 50-year bamboo seeding cycle to boost its own reproduction.In domesticating the chicken, humans took advantage of this prolific reproduction of the jungle fowl when exposed to large amounts of food.

It has been claimed (based on paleoclimatic assumptions) that chickens were domesticated in Southern China in 6000 BC. However, according to a recent study, "it is not known whether these birds made much contribution to the modern domestic fowl. Chickens from the Harappan culture of the Indus Valley (2500-2100 BC), in what today is Pakistan, may have been the main source of diffusion throughout the world." A northern road spread the chicken to the Tarim basin of central Asia. The chicken reached Europe (Romania, Turkey, Greece, Ukraine) about 3000 BC.Introduction into Western Europe came far later, about the 1st millennium BC. Phoenicians spread chickens along the Mediterranean coasts, to Iberia. Breeding increased under the Roman Empire, and was reduced in the Middle Ages. Middle East traces of chicken go back to a little earlier than 2000 BC, in Syria; chicken went southward only in the 1st millennium BC. The chicken reached Egypt for purposes of cock fighting about 1400 BC, and became widely bred only in Ptolemaic Egypt (about 300 BC).Little is known about the chicken's introduction into Africa. Three possible routes of introduction in about the early first millennium AD could have been through the Egyptian Nile Valley, the East Africa Roman-Greek or Indian trade, or from Carthage and the Berbers, across the Sahara. The earliest known remains are from Mali, Nubia, East Coast, and South Africa and date back to the middle of the first millennium AD. Domestic chicken in the Americas before Western conquest is still an ongoing discussion, but blue-egged chickens, found only in the Americas and Asia, suggest an Asian origin for early American chickens.

A lack of data from Thailand, Russia, the Indian subcontinent, Southeast Asia and Sub-Saharan Africa makes it difficult to lay out a clear map of the spread of chickens in these areas; better description and genetic analysis of local breeds threatened by extinction may also help with research into this area.

South America

An unusual variety of chicken that has its origins in South America is the araucana, bred in southern Chile by the Mapuche people. Araucanas, some of which are tailless and some of which have tufts of feathers around their ears, lay blue-green eggs. It has long been suggested that they pre-date the arrival of European chickens brought by the Spanish and are evidence of pre-Columbian trans-Pacific contacts between Asian or Pacific Oceanic peoples, particularly the Polynesians, and South America. In 2007, an international team of researchers reported the results of analysis of chicken bones found on the Arauco Peninsula in south-central Chile. Radiocarbon dating suggested that the chickens were Pre-Columbian, and DNA analysis showed that they were related to prehistoric populations of chickens in Polynesia. These results appeared to confirm that the chickens came from Polynesia and that there were transpacific contacts between Polynesia and South America before Columbus's arrival in the Americas.

However, a later report looking at the same specimens concluded:

A published, apparently pre-Columbian, Chilean specimen and six pre-European Polynesian specimens also cluster with the same European/Indian subcontinental/Southeast Asian sequences, providing no support for a Polynesian introduction of chickens to South America. In contrast, sequences from two archaeological sites on Easter Island group with an uncommon haplogroup from Indonesia, Japan, and China and may represent a genetic signature of an early Polynesian dispersal. Modeling of the potential marine carbon contribution to the Chilean archaeological specimen casts further doubt on claims for pre-Columbian chickens, and definitive proof will require further analyses of ancient DNA sequences and radiocarbon and stable isotope data from archaeological excavations within both Chile and Polynesia.

Farming

More than 50 billion chickens are reared annually as a source of food, for both their meat and their eggs.

The vast majority of poultry are raised in factory farms. According to the Worldwatch Institute, 74 percent of the world's poultry meat and 68 percent of eggs are produced this way. One alternative to intensive poultry farming is free range farming.

Friction between these two main methods has led to long-term issues of ethical consumerism. Opponents of intensive farming argue that it harms the environment, creates human health risks and is inhumane. Advocates of intensive farming say that their highly efficient systems save land and food resources owing to increased productivity, and that the animals are looked after in state-of-the-art environmentally controlled facilities.

In part due to the conditions on intensive poultry farms and recent recalls of large quantities of eggs, there is a growing movement for small-scale micro-flocks or 'backyard chickens'. This involves keeping small numbers of hens (usually no more than a dozen) in suburban or urban residential areas to control bugs, to utilize chicken waste as fertilizer in small gardens, and of course for the high-quality eggs and meat that are produced.

Reared for meat

Chickens farmed for meat are called broiler chickens. Chickens will naturally live for 6 or more years, but broiler chickens typically take less than 6 weeks to reach slaughter size.A free range or organic meat chicken will usually be slaughtered at about 14 weeks of age.

Reared for eggs

Chickens farmed for eggs are called egg-laying hens. In total, the UK alone consumes over 29 million eggs per day. Some hens breeds can produce over 300 eggs per year, with "the highest authenticated rate of egg laying being 371 eggs in 364 days". After 12 months of laying, the commercial hen's egg-laying ability starts to decline to the point where the flock is unviable. Hens, particularly from battery cage systems, are sometimes infirm or have lost a significant amount of their feathers, and their life expectancy has been reduced from around 7 years to less than 2 years. In the UK and Europe, laying hens are then slaughtered and used in processed foods or sold as "soup hens".In some other countries, flocks are sometimes force moulted, rather than being slaughtered, to reinvigorate egg-laying. This involves complete withdrawal of food (and sometimes water) for 7–14 daysor sufficiently long to cause a body weight loss of 25 to 35%, or up to 28 days under experimental conditions.This stimulates the hen to lose her feathers, but also reinvigorates egg-production. Some flocks may be force moulted several times. In 2003, more than 75% of all flocks were moulted in the US.

Artificial incubation

Incubation can successfully occur artificially in machines that provide the correct, controlled environment for the developing chick.The average incubation period for chickens is 21 days but may depend on the temperature and humidity in the incubator. Temperature regulation is the most critical factor for a successful hatch. Variations of more than 1 °C (1.8 °F) from the optimum temperature of 37.5 °C (99.5 °F) will reduce hatch rates. Humidity is also important because the rate at which eggs lose water by evaporation depends on the ambient relative humidity. Evaporation can be assessed by candling, to view the size of the air sac, or by measuring weight loss. Relative humidity should be increased to around 70% in the last three days of incubation to keep the membrane around the hatching chick from drying out after the chick cracks the shell. Lower humidity is usual in the first 18 days to ensure adequate evaporation. The position of the eggs in the incubator can also influence hatch rates. For best results, eggs should be placed with the pointed ends down and turned regularly (at least three times per day) until one to three days before hatching. If the eggs aren't turned, the embryo inside may stick to the shell and may hatch with physical defects. Adequate ventilation is necessary to provide the embryo with oxygen. Older eggs require increased ventilation.

Many commercial incubators are industrial-sized with shelves holding tens of thousands of eggs at a time, with rotation of the eggs a fully automated process. Home incubators are boxes holding from 6 to 75 eggs; they are usually electrically powered, but in the past some were heated with an oil or paraffin lamp.

As pets

Main article: Chickens as pets

Chickens are sometimes kept as pets and can be tamed by hand feeding, but roosters can sometimes become aggressive and noisy, although aggression can be curbed with proper handling. Some have advised against keeping them around very young children. Certain breeds, however, such as silkies and many bantam varieties are generally docile and are often recommended as good pets around children with disabilities.Some people find chickens' behaviour entertaining and educational.

Diseases and ailments

Chickens are susceptible to several parasites, including lice, mites, ticks, fleas, and intestinal worms, as well as other diseases. Despite the name, they are not affected by chickenpox, which is generally restricted to humans.

As food

Main article: Chicken (food)

The meat of the chicken, also called "chicken", is a type of poultry meat. Because of its relatively low cost, chicken is one of the most used meats in the world. Nearly all parts of the bird can be used for food, and the meat can be cooked in many different ways. Popular chicken dishes include roasted chicken, fried chicken, chicken soup, Buffalo wings, tandoori chicken, butter chicken, and chicken rice. Chicken is also a staple of many fast food restaurants.

Eggs

Main articles: Egg (food) and List of egg dishes

In 2000, there were 50.4 million tons of eggs produced in the world (Executive guide to world poultry trends, 2001) and an estimated 53.4 million tons of table eggs were produced during 2002. In 2009, an estimated 62.1 million metric tons of eggs were produced worldwide from a total laying flock of approximately 6.4 billion hens.

Chicken eggs are widely used in many types of dishes, both sweet and savory, including many baked goods. Eggs can be scrambled, fried, hard-boiled, soft-boiled, pickled, and poached. The albumen, or egg white, contains protein but little or no fat, and can be used in cooking separately from the yolk. Egg whites may be aerated or whipped to a light, fluffy consistency and are often used in desserts such as meringues and mousse. Ground egg shells are sometimes used as a food additive to deliver calcium. Hens do not need a male to produce eggs, only to fertilize them. A flock containing only females will still produce eggs; however, the eggs will all be infertile.

Horse

The horse (Equus ferus caballus) is one of two extant subspecies of Equus ferus. It is an odd-toed ungulate mammal belonging to the taxonomic family Equidae. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, Hyracotherium, into the large, single-toed animal of today. Humans began to domesticate horses around 4000 BC, and their domestication is believed to have been widespread by 3000 BC. Horses in the subspecies caballus are domesticated, although some domesticated populations live in the wild as feral horses. These feral populations are not true wild horses, as this term is used to describe horses that have never been domesticated, such as the endangered Przewalski's horse, a separate subspecies, and the only remaining true wild horse. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior.

Horses' anatomy enables them to make use of speed to escape predators and they have a well-developed sense of balance and a strong fight-or-flight response. Related to this need to flee from predators in the wild is an unusual trait: horses are able to sleep both standing up and lying down. Female horses, called mares, carry their young for approximately 11 months, and a young horse, called a foal, can stand and run shortly following birth. Most domesticated horses begin training under saddle or in harness between the ages of two and four. They reach full adult development by age five, and have an average lifespan of between 25 and 30 years.

Horse breeds are loosely divided into three categories based on general temperament: spirited "hot bloods" with speed and endurance; "cold bloods", such as draft horses and some ponies, suitable for slow, heavy work; and "warmbloods", developed from crosses between hot bloods and cold bloods, often focusing on creating breeds for specific riding purposes, particularly in Europe. There are more than 300 breeds of horse in the world today, developed for many different uses.

Horses and humans interact in a wide variety of sport competitions and non-competitive recreational pursuits, as well as in working activities such as police work, agriculture, entertainment, and therapy. Horses were historically used in warfare, from which a wide variety of riding and driving techniques developed, using many different styles of equipment and methods of control. Many products are derived from horses, including meat, milk, hide, hair, bone, and pharmaceuticals extracted from the urine of pregnant mares. Humans provide domesticated horses with food, water and shelter, as well as attention from specialists such as veterinarians and farriers.

>> Biology
Specific terms and specialized language are used to describe equine anatomy, different life stages, colors and breeds.

Lifespan and life stages
Depending on breed, management and environment, the modern domestic horse has a life expectancy of 25 to 30 years. Uncommonly, a few animals live into their 40s and, occasionally, beyond. The oldest verifiable record was "Old Billy", a 19th-century horse that lived to the age of 62. In modern times, Sugar Puff, who had been listed in Guinness World Records as the world's oldest living pony, died in 2007 at age 56.

Regardless of a horse or pony's actual birth date, for most competition purposes a year is added to its age each January 1 of each year in the Northern Hemisphere and each August 1 in the Southern Hemisphere.The exception is in endurance riding, where the minimum age to compete is based on the animal's actual calendar age.

The following terminology is used to describe horses of various ages:

Colt: A male horse under the age of four. A common terminology error is to call any young horse a "colt", when the term actually only refers to young male horses.
Filly: A female horse under the age of four.
Foal: A horse of either sex less than one year old. A nursing foal is sometimes called a suckling and a foal that has been weaned is called a weanling. Most domesticated foals are weaned at five to seven months of age, although foals can be weaned at four months with no adverse physical effects.
Gelding: A castrated male horse of any age
Mare: A female horse four years old and older.
Stallion: A non-castrated male horse four years oldand older. The term "horse" is sometimes used colloquially to refer specifically to a stallion.
Yearling: A horse of either sex that is between one and two years old.
In horse racing, these definitions may differ: For example, in the British Isles, Thoroughbred horse racing defines colts and fillies as less than five years old.However, Australian Thoroughbred racing defines colts and fillies as less than four years old.

Size and measurement
The height of horses is usually measured at the highest point of the withers, where the neck meets the back.This point is used because it is a stable point of the anatomy, unlike the head or neck, which move up and down in relation to the body of the horse.

In English-speaking countries, the height of horses is often stated in units of hands and inches: one hand is equal to 4 inches (101.6 mm). The height is expressed as the number of full hands, followed by a point, then the number of additional inches, and ending with the abbreviation "h" or "hh" (for "hands high"). Thus, a horse described as "15.2 h" is 15 hands plus 2 inches, for a total of 62 inches (157.5 cm) in height.
The size of horses varies by breed, but also is influenced by nutrition. Light riding horses usually range in height from 14 to 16 hands (56 to 64 inches, 142 to 163 cm) and can weigh from 380 to 550 kilograms (840 to 1,210 lb). Larger riding horses usually start at about 15.2 hands (62 inches, 157 cm) and often are as tall as 17 hands (68 inches, 173 cm), weighing from 500 to 600 kilograms (1,100 to 1,320 lb). Heavy or draft horses are usually at least 16 hands (64 inches, 163 cm) high and can be as tall as 18 hands (72 inches, 183 cm) high. They can weigh from about 700 to 1,000 kilograms (1,540 to 2,200 lb).

The largest horse in recorded history was probably a Shire horse named Mammoth, who was born in 1848. He stood 21.2 1⁄2 hands (86.5 inches, 220 cm) high and his peak weight was estimated at 1,500 kilograms (3,300 lb). The current record holder for the world's smallest horse is Thumbelina, a fully mature miniature horse affected by dwarfism. She is 17 in (43 cm) tall and weighs 57 lb (26 kg).

Ponies
Main article: Pony
Ponies are taxonomically the same animals as horses. The distinction between a horse and pony is commonly drawn on the basis of height, especially for competition purposes. However, height alone is not dispositive; the difference between horses and ponies may also include aspects of phenotype, including conformation and temperament.

The traditional standard for height of a horse or a pony at maturity is 14.2 hands (58 inches, 147 cm). An animal 14.2 h or over is usually considered to be a horse and one less than 14.2 h a pony, but there are many exceptions to the traditional standard. In Australia, ponies are considered to be those under 14 hands (56 inches, 142 cm), For competition in the Western division of the United States Equestrian Federation, the cutoff is 14.1 hands (57 inches, 145 cm) The International Federation for Equestrian Sports, the world governing body for horse sport, uses metric measurements and defines a pony as being any horse measuring less than 148 centimetres (58.27 in) at the withers without shoes, which is just over 14.2 h, and 149 centimetres (58.66 in), or just over 14.2½ h, with shoes.

Height is not the sole criterion for distinguishing horses from ponies. Breed registries for horses that typically produce individuals both under and over 14.2 h consider all animals of that breed to be horses regardless of their height. Conversely, some pony breeds may have features in common with horses, and individual animals may occasionally mature at over 14.2 h, but are still considered to be ponies.

Ponies often exhibit thicker manes, tails, and overall coat. They also have proportionally shorter legs, wider barrels, heavier bone, shorter and thicker necks, and short heads with broad foreheads. They may have calmer temperaments than horses and also a high level of equine intelligence that may or may not be used to cooperate with human handlers.Small size, by itself, is not an exclusive determinant. For example, the Shetland pony which averages 10 hands (40 inches, 102 cm), is considered a pony. Conversely, breeds such as the Falabella and other miniature horses, which can be no taller than 30 inches (76 cm), are classified by their registries as very small horses, not ponies.

Genetics
Horses have 64 chromosomes. The horse genome was sequenced in 2007. It contains 2.7 billion DNA base pairs, which is larger than the dog genome, but smaller than the human genome or the bovine genome. The map is available to researchers.

Colors and markings
Horses exhibit a diverse array of coat colors and distinctive markings, described by a specialized vocabulary. Often, a horse is classified first by its coat color, before breed or sex. Horses of the same color may be distinguished from one another by white markings, which, along with various spotting patterns, are inherited separately from coat color.

Many genes that create horse coat colors and patterns have been identified. Current genetic tests can identify at least 13 different alleles influencing coat color,and research continues to discover new genes linked to specific traits. The basic coat colors of chestnut and black are determined by the gene controlled by the Melanocortin 1 receptor, also known as the "extension gene" or "red factor," as its recessive form is "red" (chestnut) and its dominant form is black. Additional genes control suppression of black color to point coloration that results in a bay, spotting patterns such as pinto or leopard, dilution genes such as palomino or dun, as well as graying, and all the other factors that create the many possible coat colors found in horses.

Horses which have a white coat color are often mislabeled; a horse that looks "white" is usually a middle-aged or older gray. Grays are born a darker shade, get lighter as they age, but usually keep black skin underneath their white hair coat (with the exception of pink skin under white markings). The only horses properly called white are born with a predominantly white hair coat and pink skin, a fairly rare occurrence.Different and unrelated genetic factors can produce white coat colors in horses, including several different alleles of dominant white and the sabino-1 gene. However, there are no "albino" horses, defined as having both pink skin and red eyes.

Reproduction and development
Main article: Horse breeding
Gestation lasts approximately 340 days, with an average range 320–370 days, and usually results in one foal; twins are rare. Horses are a precocial species, and foals are capable of standing and running within a short time following birth.Foals are usually born in the spring. The estrous cycle of a mare occurs roughly every 19–22 days and occurs from early spring into autumn. Most mares enter an anestrus period during the winter and thus do not cycle in this period. Foals are generally weaned from their mothers between four and six months of age.

Horses, particularly colts, sometimes are physically capable of reproduction at about 18 months, but domesticated horses are rarely allowed to breed before the age of three, especially females.Horses four years old are considered mature, although the skeleton normally continues to develop until the age of six; maturation also depends on the horse's size, breed, sex, and quality of care. Larger horses have larger bones; therefore, not only do the bones take longer to form bone tissue, but the epiphyseal plates are larger and take longer to convert from cartilage to bone. These plates convert after the other parts of the bones, and are crucial to development.

Depending on maturity, breed, and work expected, horses are usually put under saddle and trained to be ridden between the ages of two and four. Although Thoroughbred race horses are put on the track as young as the age of two in some countries, horses specifically bred for sports such as dressage are generally not put under saddle until they are three or four years old, because their bones and muscles are not solidly developed. For endurance riding competition, horses are not deemed mature enough to compete until they are a full 60 calendar months (five years) old.

Anatomy
Main articles: Equine anatomy, Muscular system of the horse, Respiratory system of the horse and Circulatory system of the horse

Skeletal system
The horse skeleton averages 205 bones. A significant difference between the horse skeleton and that of a human is the lack of a collarbone—the horse's forelimbs are attached to the spinal column by a powerful set of muscles, tendons, and ligaments that attach the shoulder blade to the torso. The horse's legs and hooves are also unique structures. Their leg bones are proportioned differently from those of a human. For example, the body part that is called a horse's "knee" is actually made up of the carpal bones that correspond to the human wrist. Similarly, the hock contains bones equivalent to those in the human ankle and heel. The lower leg bones of a horse correspond to the bones of the human hand or foot, and the fetlock (incorrectly called the "ankle") is actually the proximal sesamoid bones between the cannon bones (a single equivalent to the human metacarpal or metatarsal bones) and the proximal phalanges, located where one finds the "knuckles" of a human. A horse also has no muscles in its legs below the knees and hocks, only skin, hair, bone, tendons, ligaments, cartilage, and the assorted specialized tissues that make up the hoof.

Hooves
Main articles: Horse hoof, Horseshoe and Farrier
The critical importance of the feet and legs is summed up by the traditional adage, "no foot, no horse". The horse hoof begins with the distal phalanges, the equivalent of the human fingertip or tip of the toe, surrounded by cartilage and other specialized, blood-rich soft tissues such as the laminae. The exterior hoof wall and horn of the sole is made of keratin, the same material as a human fingernail. The end result is that a horse, weighing on average 500 kilograms (1,100 lb),travels on the same bones as would a human on tiptoe. For the protection of the hoof under certain conditions, some horses have horseshoes placed on their feet by a professional farrier. The hoof continually grows, and in most domesticated horses needs to be trimmed (and horseshoes reset, if used) every five to eight weeks, though the hooves of horses in the wild wear down and regrow at a rate suitable for their terrain.

Teeth
Main article: Horse teeth
Horses are adapted to grazing. In an adult horse, there are 12 incisors at the front of the mouth, adapted to biting off the grass or other vegetation. There are 24 teeth adapted for chewing, the premolars and molars, at the back of the mouth. Stallions and geldings have four additional teeth just behind the incisors, a type of canine teeth called "tushes". Some horses, both male and female, will also develop one to four very small vestigial teeth in front of the molars, known as "wolf" teeth, which are generally removed because they can interfere with the bit. There is an empty interdental space between the incisors and the molars where the bit rests directly on the gums, or "bars" of the horse's mouth when the horse is bridled.

An estimate of a horse's age can be made from looking at its teeth. The teeth continue to erupt throughout life and are worn down by grazing. Therefore, the incisors show changes as the horse ages; they develop a distinct wear pattern, changes in tooth shape, and changes in the angle at which the chewing surfaces meet. This allows a very rough estimate of a horse's age, although diet and veterinary care can also affect the rate of tooth wear.

Digestion
Main articles: Equine digestive system and Equine nutrition
Horses are herbivores with a digestive system adapted to a forage diet of grasses and other plant material, consumed steadily throughout the day. Therefore, compared to humans, they have a relatively small stomach but very long intestines to facilitate a steady flow of nutrients. A 450-kilogram (990 lb) horse will eat 7 to 11 kilograms (15 to 24 lb) of food per day and, under normal use, drink 38 to 45 litres (8.4 to 9.9 imp gal; 10 to 12 US gal) of water. Horses are not ruminants, they have only one stomach, like humans, but unlike humans, they can utilize cellulose, a major component of grass. Horses are hindgut fermenters, Cellulose fermentation by symbiotic bacteria occurs in the cecum, or "water gut", which food goes through before reaching the large intestine. Horses cannot vomit, so digestion problems can quickly cause colic, a leading cause of death.

Senses
The horses' senses are based on their status as prey animals, where they must be aware of their surroundings at all times. They have the largest eyes of any land mammal, and are lateral-eyed, meaning that their eyes are positioned on the sides of their heads. This means that horses have a range of vision of more than 350°, with approximately 65° of this being binocular vision and the remaining 285° monocular vision.Horses have excellent day and night vision, but they have two-color, or dichromatic vision; their color vision is somewhat like red-green color blindness in humans, where certain colors, especially red and related colors, appear as a shade of green.

Their sense of smell, while much better than that of humans, is not quite as good as that of a dog. It is believed to play a key role in the social interactions of horses as well as detecting other key scents in the environment. Horses have two olfactory centers. The first system is in the nostrils and nasal cavity, which analyze a wide range of odors. The second, located under the nasal cavity, are the Vomeronasal organs, also called Jacobson's organs. These have a separate nerve pathway to the brain and appear to primarily analyze pheromones.

A horse's hearing is good, and the pinna of each ear can rotate up to 180°, giving the potential for 360° hearing without having to move the head. Noise impacts the behavior of horses and certain kinds of noise may contribute to stress: A 2013 study in the UK indicated that stabled horses were calmest in a quiet setting, or if listening to country or classical music, but displayed signs of nervousness when listening to jazz or rock music. This study also recommended keeping music under a volume of 21 decibels.An Australian study found that stabled racehorses listening to talk radio had a higher rate of gastric ulcers than horses listening to music, and racehorses stabled where a radio was played had a higher overall rate of ulceration than horses stabled where there was no radio playing.

Horses have a great sense of balance, due partly to their ability to feel their footing and partly to highly developed proprioception—the unconscious sense of where the body and limbs are at all times. A horse's sense of touch is well developed. The most sensitive areas are around the eyes, ears, and nose.Horses are able to sense contact as subtle as an insect landing anywhere on the body.

Horses have an advanced sense of taste, which allows them to sort through fodder and choose what they would most like to eat, and their prehensile lips can easily sort even small grains. Horses generally will not eat poisonous plants, however, there are exceptions; horses will occasionally eat toxic amounts of poisonous plants even when there is adequate healthy food.
Movement
All horses move naturally with four basic gaits: the four-beat walk, which averages 6.4 kilometres per hour (4.0 mph); the two-beat trot or jog at 13 to 19 kilometres per hour (8.1 to 11.8 mph) (faster for harness racing horses); the canter or lope, a three-beat gait that is 19 to 24 kilometres per hour (12 to 15 mph); and the gallop. The gallop averages 40 to 48 kilometres per hour (25 to 30 mph), but the world record for a horse galloping over a short, sprint distance is 88 kilometres per hour (55 mph).Besides these basic gaits, some horses perform a two-beat pace, instead of the trot. There also are several four-beat "ambling" gaits that are approximately the speed of a trot or pace, though smoother to ride. These include the lateral rack, running walk, and tölt as well as the diagonal fox trot. Ambling gaits are often genetic in some breeds, known collectively as gaited horses. Often, gaited horses replace the trot with one of the ambling gaits.

Behavior
Main articles: Horse behavior and Stable vices
Horses are prey animals with a strong fight-or-flight response. Their first reaction to threat is to startle and usually flee, although they will stand their ground and defend themselves when flight is impossible or if their young are threatened. They also tend to be curious; when startled, they will often hesitate an instant to ascertain the cause of their fright, and may not always flee from something that they perceive as non-threatening. Most light horse riding breeds were developed for speed, agility, alertness and endurance; natural qualities that extend from their wild ancestors. However, through selective breeding, some breeds of horses are quite docile, particularly certain draft horses.

Horses are herd animals, with a clear hierarchy of rank, led by a dominant individual, usually a mare. They are also social creatures that are able to form companionship attachments to their own species and to other animals, including humans. They communicate in various ways, including vocalizations such as nickering or whinnying, mutual grooming, and body language. Many horses will become difficult to manage if they are isolated, but with training, horses can learn to accept a human as a companion, and thus be comfortable away from other horses. However, when confined with insufficient companionship, exercise, or stimulation, individuals may develop stable vices, an assortment of bad habits, mostly stereotypies of psychological origin, that include wood chewing, wall kicking, "weaving" (rocking back and forth), and other problems.
Intelligence and learning
Studies have indicated that horses perform a number of cognitive tasks on a daily basis, meeting mental challenges that include food procurement and identification of individuals within a social system. They also have good spatial discrimination abilities. Studies have assessed equine intelligence in areas such as problem solving, speed of learning, and memory. Horses excel at simple learning, but also are able to use more advanced cognitive abilities that involve categorization and concept learning. They can learn using habituation, desensitization, classical conditioning, and operant conditioning, and positive and negative reinforcement.One study has indicated that horses can differentiate between "more or less" if the quantity involved is less than four.

Domesticated horses may face greater mental challenges than wild horses, because they live in artificial environments that prevent instinctive behavior whilst also learning tasks that are not natural.Horses are animals of habit that respond well to regimentation, and respond best when the same routines and techniques are used consistently. One trainer believes that "intelligent" horses are reflections of intelligent trainers who effectively use response conditioning techniques and positive reinforcement to train in the style that best fits with an individual animal's natural inclinations.

Temperament
Horses are mammals, and as such are warm-blooded, or endothermic creatures, as opposed to cold-blooded, or poikilothermic animals. However, these words have developed a separate meaning in the context of equine terminology, used to describe temperament, not body temperature. For example, the "hot-bloods", such as many race horses, exhibit more sensitivity and energy, while the "cold-bloods", such as most draft breeds, are quieter and calmer. Sometimes "hot-bloods" are classified as "light horses" or "riding horses", with the "cold-bloods" classified as "draft horses" or "work horses".
"Hot blooded" breeds include "oriental horses" such as the Akhal-Teke, Arabian horse, Barb and now-extinct Turkoman horse, as well as the Thoroughbred, a breed developed in England from the older oriental breeds. Hot bloods tend to be spirited, bold, and learn quickly. They are bred for agility and speed.They tend to be physically refined—thin-skinned, slim, and long-legged.The original oriental breeds were brought to Europe from the Middle East and North Africa when European breeders wished to infuse these traits into racing and light cavalry horses.

Muscular, heavy draft horses are known as "cold bloods", as they are bred not only for strength, but also to have the calm, patient temperament needed to pull a plow or a heavy carriage full of people. They are sometimes nicknamed "gentle giants".Well-known draft breeds include the Belgian and the Clydesdale.Some, like the Percheron, are lighter and livelier, developed to pull carriages or to plow large fields in drier climates. Others, such as the Shire, are slower and more powerful, bred to plow fields with heavy, clay-based soils.The cold-blooded group also includes some pony breeds.

"Warmblood" breeds, such as the Trakehner or Hanoverian, developed when European carriage and war horses were crossed with Arabians or Thoroughbreds, producing a riding horse with more refinement than a draft horse, but greater size and milder temperament than a lighter breed.Certain pony breeds with warmblood characteristics have been developed for smaller riders.Warmbloods are considered a "light horse" or "riding horse".

Today, the term "Warmblood" refers to a specific subset of sport horse breeds that are used for competition in dressage and show jumping.Strictly speaking, the term "warm blood" refers to any cross between cold-blooded and hot-blooded breeds.Examples include breeds such as the Irish Draught or the Cleveland Bay. The term was once used to refer to breeds of light riding horse other than Thoroughbreds or Arabians, such as the Morgan horse.

Sleep patterns
See also: Horse sleep patterns and Sleep in non-humans
Horses are able to sleep both standing up and lying down. In an adaptation from life in the wild, horses are able to enter light sleep by using a "stay apparatus" in their legs, allowing them to doze without collapsing. Horses sleep better when in groups because some animals will sleep while others stand guard to watch for predators. A horse kept alone will not sleep well because its instincts are to keep a constant eye out for danger.

Unlike humans, horses do not sleep in a solid, unbroken period of time, but take many short periods of rest. Horses spend four to fifteen hours a day in standing rest, and from a few minutes to several hours lying down. Total sleep time in a 24-hour period may range from several minutes to a couple of hours, mostly in short intervals of about 15 minutes each.The average sleep time of a domestic horse is said to be 2.9 hours per day.

Horses must lie down to reach REM sleep. They only have to lie down for an hour or two every few days to meet their minimum REM sleep requirements. However, if a horse is never allowed to lie down, after several days it will become sleep-deprived, and in rare cases may suddenly collapse as it involuntarily slips into REM sleep while still standing.This condition differs from narcolepsy, although horses may also suffer from that disorder.

>> Taxonomy and evolution
The horse adapted to survive in areas of wide-open terrain with sparse vegetation, surviving in an ecosystem where other large grazing animals, especially ruminants, could not.Horses and other equids are odd-toed ungulates of the order Perissodactyla, a group of mammals that was dominant during the Tertiary period. In the past, this order contained 14 families, but only three—Equidae (the horse and related species), the tapir, and the rhinoceros—have survived to the present day.

The earliest known member of the Equidae family was the Hyracotherium, which lived between 45 and 55 million years ago, during the Eocene period. It had 4 toes on each front foot, and 3 toes on each back foot.The extra toe on the front feet soon disappeared with the Mesohippus, which lived 32 to 37 million years ago. Over time, the extra side toes shrank in size until they vanished. All that remains of them in modern horses is a set of small vestigial bones on the leg below the knee,known informally as splint bones.Their legs also lengthened as their toes disappeared until they were a hooved animal capable of running at great speed. By about 5 million years ago, the modern Equus had evolved.Equid teeth also evolved from browsing on soft, tropical plants to adapt to browsing of drier plant material, then to grazing of tougher plains grasses. Thus proto-horses changed from leaf-eating forest-dwellers to grass-eating inhabitants of semi-arid regions worldwide, including the steppes of Eurasia and the Great Plains of North America.

By about 15,000 years ago, Equus ferus was a widespread holarctic species. Horse bones from this time period, the late Pleistocene, are found in Europe, Eurasia, Beringia, and North America.Yet between 10,000 and 7,600 years ago, the horse became extinct in North America and rare elsewhere.The reasons for this extinction are not fully known, but one theory notes that extinction in North America paralleled human arrival.Another theory points to climate change, noting that approximately 12,500 years ago, the grasses characteristic of a steppe ecosystem gave way to shrub tundra, which was covered with unpalatable plants.